TCS: Phylogenetic network estimation using statistical parsimony
© 2000-2005 Mark Clement, Jacob Derington, Steve Woolley and David Posada
Current version is 1.21
TCS is a Java computer program to estimate gene genealogies including multifurcations and/or reticulations (i.e. networks). The network estimation implemented in TCS is also known as Statistical Parsimony, which is described in Templeton, A. R., K. A. Crandall and C. F. Sing. 1992. A cladistic analysis of phenotypic associations with haplotypes inferred from restriction endonuclease mapping and DNA sequence data. III. Cladogram estimation. Genetics 132:619-633. For a review on networks and instraspecific genealogies you may read Posada D and Crandall KA. 2001. Trends in Ecology and Evolution 16 (1): 37-45
A jar file is provided that should run in any OS (macintosh, windows, unix-like) with a java virtual machine installed.
Clement M, Posada D and Crandall K. 2000. TCS: a computer program to estimate gene genealogies. Molecular Ecology 9(10): 1657-1660
Disclaimer: These programs are free software; you can redistribute it and/or modify it under the terms of the GNU General Public License as published by the Free Software Foundation; either version 2 of the License, or (at your option) any later version. These programs are distributed in the hope that it will be useful, but WITHOUT ANY WARRANTY; without even the implied warranty of MERCHANTABILITY or FITNESS FOR A PARTICULAR PURPOSE. See the GNU General Public License for more details.
Correspondence about this program should be addressed to David Posada at
This document last modified Thursday June 06, 2013