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GEODIS: A tool to perform the nested clade phylogeographic analysis (NCPA)

© 1999-2009 David Posada and Alan Templeton

Current version is 2.6 (28 April 2009)

GEODIS is a program for the calculation of the statistics and associated P-values for the nested clade analysis (NCA) developed by Templeton and collaborators. In the NCA, population structure can be separated from population history when it is assessed through rigorous and objective statistical tests using the temporal information contained in the a genealogy.

NEW! The last version (2.6) includes the Dunn-Sidak correction for multiple tests.

GeoDis inference key
NEW! Make sure you have the latest inference key: geodis inference key (January 06, 2011).

Operative systems
A jar file is provided that should run in any OS (macintosh, windows, unix-like) with a java virtual machine installed.

Posada D, Crandall KA and Templeton AR. 2000. GeoDis: A program for the Cladistic Nested Analysis of the Geographical Distribution of Genetic Haplotypes. Molecular Ecology 9(4):487-488.

GeoDis registration form

Before download this program, I ask users to register. This allows me to inform about bugs and updates in a more efficient way. Indeed, users will be able to unregister at any time, and email addresses will not be used for any purpose other than to inform about this program.


Any comments?

Disclaimer: These programs are free software; you can redistribute it and/or modify it under the terms of the GNU General Public License as published by the Free Software Foundation; either version 2 of the License, or (at your option) any later version. These programs are distributed in the hope that it will be useful, but WITHOUT ANY WARRANTY; without even the implied warranty of MERCHANTABILITY or FITNESS FOR A PARTICULAR PURPOSE. See the GNU General Public License for more details.

Correspondence about this program should be addressed to David Posada at

This document last modified Thursday June 06, 2013